Authors Authors and affiliations Grimme L. Horst Jeanette S. This process is experimental and the keywords may be updated as the learning algorithm improves. This is a preview of subscription content, log in to check access. Braumann Th, Weber G and Grimme LH Carotenoid and chlorophyll composition of light-harvesting and reaction centre proteins of the thylakoid membrane, Photobiochem.
Google Scholar. CrossRef Google Scholar. Brown JS and Schoch S Spectral analysis of chlorophyll-protein complexes from higher plant chloroplasts, Biochim.
Acta , — Carnegie Inst. When light energy reaches the pigment molecules, it energizes the electrons within them, and these electrons are shunted to an electron transport chain in the thylakoid membrane. Meanwhile, each chlorophyll molecule replaces its lost electron with an electron from water; this process essentially splits water molecules to produce oxygen Figure 5.
Figure 5: The light and dark reactions in the chloroplast The chloroplast is involved in both stages of photosynthesis. The light reactions take place in the thylakoid. There, water H 2 O is oxidized, and oxygen O 2 is released. The dark reactions then occur outside the thylakoid. The products of this reaction are sugar molecules and various other organic molecules necessary for cell function and metabolism.
Note that the dark reaction takes place in the stroma the aqueous fluid surrounding the stacks of thylakoids and in the cytoplasm. The thylakoids, intake of water H 2 O , and release of oxygen O 2 occur on the yellow side of the cell to indicate that these are involved in the light reactions. The carbon fixation reactions, which involve the intake of carbon dioxide CO 2 , NADPH, and ATP, and the production of sugars, fatty acids, and amino acids, occur on the blue side of the cell to indicate that these are dark reactions.
An arrow shows the movement of a water molecule from the outside to the thylakoid stack on the inside of the chloroplast. Another arrow shows light energy from the sun entering the chloroplast and reaching the thylakoid stack.
An arrow shows the release of an oxygen molecule O 2 from the thylakoid stack to the outside of the chloroplast. Once the light reactions have occurred, the light-independent or "dark" reactions take place in the chloroplast stroma. During this process, also known as carbon fixation, energy from the ATP and NADPH molecules generated by the light reactions drives a chemical pathway that uses the carbon in carbon dioxide from the atmosphere to build a three-carbon sugar called glyceraldehydephosphate G3P.
Cells then use G3P to build a wide variety of other sugars such as glucose and organic molecules. Many of these interconversions occur outside the chloroplast, following the transport of G3P from the stroma. The products of these reactions are then transported to other parts of the cell, including the mitochondria, where they are broken down to make more energy carrier molecules to satisfy the metabolic demands of the cell.
In plants, some sugar molecules are stored as sucrose or starch. This page appears in the following eBook. Aa Aa Aa. Photosynthetic Cells. What Is Photosynthesis? Why Is it Important? Figure 2. Figure 3: Structure of a chloroplast. Figure 4: Diagram of a chloroplast inside a cell, showing thylakoid stacks. Shown here is a chloroplast inside a cell, with the outer membrane OE and inner membrane IE labeled.
What Are the Steps of Photosynthesis? Figure 5: The light and dark reactions in the chloroplast. The chloroplast is involved in both stages of photosynthesis. Photosynthetic cells contain chlorophyll and other light-sensitive pigments that capture solar energy. In the presence of carbon dioxide, such cells are able to convert this solar energy into energy-rich organic molecules, such as glucose.
These cells not only drive the global carbon cycle, but they also produce much of the oxygen present in atmosphere of the Earth. Essentially, nonphotosynthetic cells use the products of photosynthesis to do the opposite of photosynthesis: break down glucose and release carbon dioxide.
Cell Biology for Seminars, Unit 1. Topic rooms within Cell Biology Close. No topic rooms are there. Or Browse Visually. Student Voices. Creature Cast. Simply Science. Green Screen. Green Science. Bio 2. The Success Code. Why Science Matters. The Beyond. Plant ChemCast. Postcards from the Universe. In eukaryotes and some prokaryotes, two photosystems exist.
The first is called photosystem II, which was named for the order of its discovery rather than for the order of the function. After the photon hits, photosystem II transfers the free electron to the first in a series of proteins inside the thylakoid membrane called the electron transport chain. As the electron passes along these proteins, energy from the electron fuels membrane pumps that actively move hydrogen ions against their concentration gradient from the stroma into the thylakoid space.
This is quite analogous to the process that occurs in the mitochondrion in which an electron transport chain pumps hydrogen ions from the mitochondrial stroma across the inner membrane and into the intermembrane space, creating an electrochemical gradient. After the energy is used, the electron is accepted by a pigment molecule in the next photosystem, which is called photosystem I Figure 5.
The energy that these molecules carry is stored in a bond that holds a single atom to the molecule. Recall that NADH was a similar molecule that carried energy in the mitochondrion from the citric acid cycle to the electron transport chain. This potential energy is harvested and stored as chemical energy in ATP through chemiosmosis, the movement of hydrogen ions down their electrochemical gradient through the transmembrane enzyme ATP synthase, just as in the mitochondrion.
The hydrogen ions are allowed to pass through the thylakoid membrane through an embedded protein complex called ATP synthase. The energy generated by the hydrogen ion stream allows ATP synthase to attach a third phosphate to ADP, which forms a molecule of ATP in a process called photophosphorylation. The flow of hydrogen ions through ATP synthase is called chemiosmosis, because the ions move from an area of high to low concentration through a semi-permeable structure.
The remaining function of the light-dependent reaction is to generate the other energy-carrier molecule, NADPH. As the electron from the electron transport chain arrives at photosystem I, it is re-energized with another photon captured by chlorophyll. Now that the solar energy is stored in energy carriers, it can be used to make a sugar molecule.
In the first part of photosynthesis, the light-dependent reaction, pigment molecules absorb energy from sunlight. The most common and abundant pigment is chlorophyll a. A photon strikes photosystem II to initiate photosynthesis. Energy travels through the electron transport chain, which pumps hydrogen ions into the thylakoid space. This forms an electrochemical gradient. The ions flow through ATP synthase from the thylakoid space into the stroma in a process called chemiosmosis to form molecules of ATP, which are used for the formation of sugar molecules in the second stage of photosynthesis.
Photosystem I absorbs a second photon, which results in the formation of an NADPH molecule, another energy carrier for the Calvin cycle reactions. Skip to content Chapter 5: Introduction to Photosynthesis. Learning Objectives By the end of this section, you will be able to: Explain how plants absorb energy from sunlight Describe how the wavelength of light affects its energy and color Describe how and where photosynthesis takes place within a plant.
Concept in Action Visit this site and click through the animation to view the process of photosynthesis within a leaf.
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